1996-2000
University of Sheffield, Western Bank, Sheffield, UK S10 2TA
PhD in Evolutionary Ecology.

1992-1995
Cambridge University, Gonville and Caius College, Cambridge, UK CB2 1TA
BA Hons in Natural Sciences: 2.1 (MA 1999)

1983-1991
Dulwich College, Dulwich Common, London, UK SE21 7LD
A level: Biology (A), Chemistry (B), Physics (B); AS level: Maths (A)